APA (7th ed.) Citation

Whittaker, R. J., & Cottee-Jones, H. E. W. (2012, September). The keystone species concept: A critical appraisal. Frontiers of Biogeography.

Chicago Style (17th ed.) Citation

Whittaker, Robert J., and Henry Eden W. Cottee-Jones. "The Keystone Species Concept: A Critical Appraisal." Frontiers of Biogeography Sep. 2012.

MLA (9th ed.) Citation

Whittaker, Robert J., and Henry Eden W. Cottee-Jones. "The Keystone Species Concept: A Critical Appraisal." Frontiers of Biogeography, Sep. 2012.