Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain

The Caledonian pinewoods of northern Scotland are a priority conservation habitat in Europe which are dominated by Scots pine (Pinus sylvestris), but varying proportions of a number of broadleaved species such as silver birch (Betula pendula) can occur in these forests. Better understanding of the d...

Full description

Bibliographic Details
Main Authors: Bill Mason, Thomas Connolly
Format: Article
Language:English
Published: Consiglio per la ricerca in agricoltura e l'analisi dell'economia agraria - Research Centre for Forestry and Wood 2016-07-01
Series:Annals of Silvicultural Research
Subjects:
Online Access:http://cra-journals.cineca.it/index.php/asr/article/view/1119
id doaj-37c0bb849aa3474d83f06cfdc44bfbb6
record_format Article
spelling doaj-37c0bb849aa3474d83f06cfdc44bfbb62020-11-24T22:32:33ZengConsiglio per la ricerca in agricoltura e l'analisi dell'economia agraria - Research Centre for Forestry and WoodAnnals of Silvicultural Research2284-354X2016-07-01401111810.12899/asr-1119911Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern BritainBill Mason0Thomas Connolly1Forest Research UKForest Research UKThe Caledonian pinewoods of northern Scotland are a priority conservation habitat in Europe which are dominated by Scots pine (Pinus sylvestris), but varying proportions of a number of broadleaved species such as silver birch (Betula pendula) can occur in these forests. Better understanding of the dynamics of mixed Scots pine-birch stands would be helpful in informing current initiatives to restore and increase the area of the pinewood ecosystem. Some evidence is provided by two experiments established in the 1960s which compared plots of pure Scots pine and pure birch with two treatments where the two species were mixed in 3:1 and 1:1 ratios. Some fifty years later, Scots pine was the more vigorous of the two species in these experiments, being both taller and significantly larger in diameter. The highest basal area was generally found in the pure Scots pine plots and the values in the mixed plots tended to be intermediate between those of the two component species. Examination of the growth in the mixed plots showed a slight, but non-significant, tendency towards overyielding. This appeared to be due to Scots pine growth being better than predicted, while that of birch was slightly less than predicted. These results suggest that in these mixtures, which are composed of two light demanding species, the main mechanism driving long-term performance is competition for light and there is little evidence of any complementary effect. These results suggest that any strategy seeking to increase the long-term representation of broadleaves such as birch in the Caledonian pinewoods will need to create discrete blocks that are large enough to withstand the competitive pressures exerted by the pine.http://cra-journals.cineca.it/index.php/asr/article/view/1119MixturesPinus sylvestris, Betula pendula, competition
collection DOAJ
language English
format Article
sources DOAJ
author Bill Mason
Thomas Connolly
spellingShingle Bill Mason
Thomas Connolly
Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain
Annals of Silvicultural Research
Mixtures
Pinus sylvestris, Betula pendula, competition
author_facet Bill Mason
Thomas Connolly
author_sort Bill Mason
title Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain
title_short Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain
title_full Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain
title_fullStr Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain
title_full_unstemmed Long-term development of experimental mixtures of Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) in northern Britain
title_sort long-term development of experimental mixtures of scots pine (pinus sylvestris l.) and silver birch (betula pendula roth.) in northern britain
publisher Consiglio per la ricerca in agricoltura e l'analisi dell'economia agraria - Research Centre for Forestry and Wood
series Annals of Silvicultural Research
issn 2284-354X
publishDate 2016-07-01
description The Caledonian pinewoods of northern Scotland are a priority conservation habitat in Europe which are dominated by Scots pine (Pinus sylvestris), but varying proportions of a number of broadleaved species such as silver birch (Betula pendula) can occur in these forests. Better understanding of the dynamics of mixed Scots pine-birch stands would be helpful in informing current initiatives to restore and increase the area of the pinewood ecosystem. Some evidence is provided by two experiments established in the 1960s which compared plots of pure Scots pine and pure birch with two treatments where the two species were mixed in 3:1 and 1:1 ratios. Some fifty years later, Scots pine was the more vigorous of the two species in these experiments, being both taller and significantly larger in diameter. The highest basal area was generally found in the pure Scots pine plots and the values in the mixed plots tended to be intermediate between those of the two component species. Examination of the growth in the mixed plots showed a slight, but non-significant, tendency towards overyielding. This appeared to be due to Scots pine growth being better than predicted, while that of birch was slightly less than predicted. These results suggest that in these mixtures, which are composed of two light demanding species, the main mechanism driving long-term performance is competition for light and there is little evidence of any complementary effect. These results suggest that any strategy seeking to increase the long-term representation of broadleaves such as birch in the Caledonian pinewoods will need to create discrete blocks that are large enough to withstand the competitive pressures exerted by the pine.
topic Mixtures
Pinus sylvestris, Betula pendula, competition
url http://cra-journals.cineca.it/index.php/asr/article/view/1119
work_keys_str_mv AT billmason longtermdevelopmentofexperimentalmixturesofscotspinepinussylvestrislandsilverbirchbetulapendularothinnorthernbritain
AT thomasconnolly longtermdevelopmentofexperimentalmixturesofscotspinepinussylvestrislandsilverbirchbetulapendularothinnorthernbritain
_version_ 1725733368030560256