Taxonomic composition of the family Trogidae (Coleoptera: Scarabaeoidea) of the Russian fauna
Glyptotrox gen. n. is described. Trox mandli Balthasar, 1931, living in the east of Russia, in China, Mongolia, Korean Peninsula and in Japan, is designated as type species of this genus. Diagnosis of the genus: small to medium size species; body oblong-oval and strongly соnvех; clypeus triangu...
Main Author: | |
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Format: | Article |
Language: | Russian |
Published: |
Russian Academy of Science. Southern Scientific Centre. Federal Research Centre
2016-07-01
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Series: | Кавказский энтомологический бюллетень |
Subjects: | |
Online Access: | http://www.ssc-ras.ru/ckfinder/userfiles/files/11_GV%20Nikolajev.pdf |
Summary: | Glyptotrox gen. n. is described. Trox mandli
Balthasar, 1931, living in the east of Russia, in China,
Mongolia, Korean Peninsula and in Japan, is designated as
type species of this genus.
Diagnosis of the genus: small to medium size species;
body oblong-oval and strongly соnvех; clypeus triangular; pronotum with anterior margin widely arcuate, lateral
margins gently rounded or sinuate, pronotal disc with six
more or less distinct, rounded or oval concavities with
limited by strongly elevated obtuse ridges; male genitalia
with narrow, elongate parameres and median lobe equipped
with more or less distinct, triangular or ear-like, subapical
processes (Figs 5a, 6a).
Currently 31 taxa (species and subspecies)
inhabiting North and Central America (Cuba), the
Eastern Palaearctic and Oriental Region are known. The
following new combinations are established: Glyptotrox
boucomonti (Paulian, 1933), comb. n., Glyptotrox
brahminus (Pittino, 1985), comb. n., Glyptotrox cambeforti
(Pittino, 1985), comb. n., Glyptotrox cambodjanus
(Pittino, 1985), comb. n., Glyptotrox dhaulagiri
(Paulus, 1972), comb. n., Glyptotrox doiinthanonensis
(Masumoto, 1996), comb. n., Glyptotrox formosanus
(Nomura, 1973), comb. n., Glyptotrox foveicollis (Harold,
1872), comb. n., Glyptotrox frontera (Vaurie, 1955), comb. n.,
Glyptotrox hamatus (Robinson, 1940), comb. n.,
Glyptotrox ineptus (Balthasar, 1931), comb. n. (= Trox
vimmeri Balthasar, 1931 synonymy by Nikolajev [1976]),
Glyptotrox insularis (Chevrolat, 1864), comb. n.,
Glyptotrox kerleyi (Masumoto, 1996), comb. n., Glyptotrox
kiuchii (Masumoto, 1996), comb. n., Glyptotrox mandli
(Balthasar, 1931), comb. n., Glyptotrox matsudai
(Ochi et Hori, 1999), comb. n., Glyptotrox mutsuensis
(Nomura, 1937), comb. n., Glyptotrox opacotuberculatus
(Motschulsky, 1860), comb. n., Glyptotrox parvisetosus
(Ochi, Kon et Bai, 2010), comb. n., Glyptotrox poringensis
(Ochi, Kon et Kawahara, 2005), comb. n., Glyptotrox
simi (Robinson, 1940), comb. n., Glyptotrox spinulosus
(Robinson, 1940), comb. n., Glyptotrox spinulosus ssp.
dentibius (Robinson, 1940), comb. n., Glyptotrox sugayai
(Masumoto et Kiuchi, 1995), comb. n., Glyptotrox
taiwanus (Masumoto, Ochi et Li, 2005), comb. n.,
Glyptotrox terrestris (Say, 1825), comb. n., Glyptotrox
tibialis (Masumoto, Ochi et Li, 2005), comb. n., Glyptotrox
uenoi (Nomura, 1961), comb. n., Glyptotrox uenoi
ssp. matsumurai (Miyake et Yamaya, 1995), comb. n.,
Glyptotrox yamayai (Nakane, 1983), comb. n., Glyptotrox
yangi (Masumoto, Ochi et Li, 2005), comb. n.
Two new subgenera in the genus Trox are described:
Granulitrox Nikolajev, subgen. n. and Niditrox
Nikolajev, subgen. n. Trox granulipennis Fairmaire, 1852
is designated as type species of the subgenus Granulitrox
Nikolajev, subgen. n. Twenty two species are included in
this subgenus: T. (Granulitrox) confluens Wollaston, 1864,
T. (Granulitrox) cribrum Gené, 1836, T. (Granulitrox)
cyrenaicus Pittino, 2011, T. (Granulitrox) elkantaraensis
Pittino, 2011, T. (Granulitrox) eximius Faldermann, 1835,
T. (Granulitrox) fabricii Reiche, 1853, T. (Granulitrox)
granulipennis Fairmaire, 1852, T. (Granulitrox) hispanicus
Harold, 1862, T. (Granulitrox) hispidus (Pontoppidan,
1763), T. (Granulitrox) iranicus Petrovitz, 1980,
T. (Granulitrox) klapperichi Pittino, 1983, T. (Granulitrox)
leonardii Pittino, 1983, T. (Granulitrox) litoralis
Pittino, 1991, T. (Granulitrox) martini Reitter, 1892,
T. (Granulitrox) mixtus Harold, 1872, T. (Granulitrox)
morticinii Pallas, 1781, T. (Granulitrox) niger P. Rossi, 1792,
T. (Granulitrox) nodulosus Harold, 1872, T. (Granulitrox)
perlatus (Goeze, 1777), T. (Granulitrox) quadrimaculatus
Ballion, 1871, T. (Granulitrox) sordidatus Balthasar, 1936,
T. (Granulitrox) transversus Reiche, 1856.
Diagnosis of the subgenus Granulitrox
Nikolajev, subgen. n.: medium size (rarely large) species;
body oblong-oval and соnvех; clypeus triangular (rounded
in large species); pronotum with lateral margins gently
rounded; scutellum triangular; elytra may have elevations
in the form of grains the bristles may be located only at
the rear boundary of the convexes on intervals between
grooves of elytrae.
Trox eversmannii Krynicki, 1832 as type species of the
subgenus Niditrox Nikolajev, subgen. n. is designated. Ten
taxa are included in the subgenus: T. (Niditrox) aequalis
Say, 1831, T. (Niditrox) affinis Robinson, 1940, T. (Niditrox)
eversmannii Krynicki, 1832, T. (Niditrox) fabricii Reiche,
1853, T. (Niditrox) koreanus J.I. Kim, 1991, T. (Niditrox)
niponensis Lewis, 1895, T. (Niditrox) nohirai Nakane, 1954,
T. (Niditrox) perrisii Fairmaire, 1868, T. (Niditrox) scaber
(Linnaeus, 1767), T. (Niditrox) zoufali Balthasar, 1931.
Diagnosis of the subgenus Niditrox Nikolajev,
subgen. n.: small to medium size species; body elongate;
clypeus triangular; lateral margins of pronotum broadly
rounded; male genitalia with narrow, elongate parameres
outstanding over the apex of median lobe (Figs 3, 4).
It is expected that molecular-genetic analyses will
be more useful than classical morphology for division of
sibling species complex similar to T. (Niditrox) scaber
(Linnaeus, 1767). |
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ISSN: | 1814-3326 1814-3326 |