| Summary: | This thesis investigates the influence of niche specialisation on floristic composition and species distributions of the family Dipterocarpaceae along topographic gradients at two contrasting lowland Mixed Dipterocarp Forest sites in Brunei Darussalam: Andulau (overlying nutrient-poor sandy soils) and Belalong (overlying nutrient-rich clay soils). Dipterocarp tree density, species richness and diversity were higher at Andulau than Belalong, and were also higher on ridges than in slopes and valleys. Randomisation tests detected significant dipterocarp species associations with site and tomography. Dipterocarp floristic composition was strongly correlated with various habitat variables at Belalong, even after accounting for distance between samples. In contrast, fewer habitat variables were correlated with dipterocarp floristic composition at Andulau, implying weaker habitat effects in this more topographically homogeneous site. In a field-based transplant experiment, <i>Dryobalanops aromatica </i>and <i>Dryobalanops rappa </i>seedlings consistently showed higher growth rates and survival in gaps than in understorey plots. Higher survival and leaf production of <i>D. aromatica </i>seedlings at Andulau than Belalong provide evidence of a habitat preference. Growth and survival of <i>D. rappa</i> seedlings were similar at the two sites, but <i>D. rappa</i> seedlings grew significantly faster in height than <i>D. aromatica </i>seedlings on slopes at both sites. In conclusion, local and landscape scale variations in edaphic and environmental resources influence dipterocarp species distributions and floristic composition, as well as dipterocarp seedling growth and survival. Thus, niche specialisation is an important mechanism in the maintenance of species coexistence at Andulau and Belalong.
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