Anthropogenic disturbance 1 Benthic ecosystems Ecosystems 1 Fiddler crab 1 Flagship species 1 Japan 1 Monitoring 1 Recreation ecology 1 Shellfish 1 South Korea 1 Surface activities 1 Surface behavior 1 Temperate regions 1 Tidal flat 1 Tidal rhythm 1 Tides 1 Wetlands 1 behavioral ecology 1 circadian rhythm 1 courtship 1 crab 1 endangered species 1 feeding 1 human activity 1 intertidal community 1 population dynamics 1 tidal flat 1 trampling 1 vulnerability 1