Arthrobotrys oligospora Article ER stress Fungistatic mechanism Multi-omics Soil fungistasis Western blotting alkalinity ammonia carbon metabolism conidium endoplasmic reticulum stress fungal spore germination fungistatic activity fungus growth fungus spore gene knockout glutamic acid metabolite multiomics nematophagous fungus nonhuman protein homeostasis protein misfolding proteome survival ubiquitination unfolded protein response upregulation